Abstract
Stem bleeding on horse chestnut (Aesculus hippocastanum) in the UK was first reported in the 1970s, when the cause was found to be a fungal pathogen known as Phytophthora (Brasier and Strouts, 1976). The same disorder had also been recognised in the USA much earlier in the 1930’s (Caroselli, 1953). Symptoms visible on affected trees include bleeding areas on their stems and sometimes on their scaffold branches. However, over the past four or five years the number of reports of horse chestnut trees with 'bleeding cankers' has increased markedly. Closer investigation of the bleeding cankers on horse chestnut has revealed that Phytophthora is no longer the primary causal agent. Instead there is accumulating evidence that a different pathogen, probably a bacterium, is responsible for the increase in these symptoms on horse chestnut.
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Claims 3
Claim · model-written summary
Reports of horse chestnut bleeding cankers increased markedly over four or five years.
Verbatim
e0“over the past four or five years the number of reports of horse chestnut trees with 'bleeding cankers' has increased markedly.”
Claim · model-written summary
Phytophthora was no longer the primary causal agent of horse chestnut bleeding cankers.
- Taxon
- horse chestnute0
- Interactor
- Phytophthora
Phytophthorae0pathogen - Outcome
- Phytophthora is no longer the primary causal agente0
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“Closer investigation of the bleeding cankers on horse chestnut has revealed that Phytophthora is no longer the primary causal agent.”
Claim · model-written summary
Evidence suggests that a different pathogen, probably a bacterium, accounts for the increase in horse chestnut symptoms.
Verbatim
e0“there is accumulating evidence that a different pathogen, probably a bacterium, is responsible for the increase in these symptoms on horse chestnut.”