Abstract
In this paper we present the flora of Bili Skeli (White Cliffs) Massif. This site is the southernmost group of limestone cliffs in the Ukrainian Carpathians, situated in the montane forest belt in the headwaters of the Bilyi Cheremosh River (Verkhovynskyi National Nature Park, the Chyvchyny Mts.). These cliffs harbour rare calcareous saxicolous species, e.g. Arabis hornungiana, Carduus defloratus subsp. glaucus, Cystopteris alpina, Epipactis atrorubens, E. helleborine, Goodyera repens, Saussurea discolor, Saxifraga luteoviridis and Thymus pulcherrimus, many of them included in the Red Data Book of Ukraine (2009). Their occurrence was analysed in their ecological and conservation context. A map of the site is provided as well as geographical coordinates of the major cliffs. The occurrence of obligatory calcicole rare species is determined by the limestone bedrock, which is infrequent in the Ukrainian Carpathians. The rarity of the habitat type makes the site one of the few disjunct environmental islands of calcareous flora in the region. Some of the listed species are narrow-range and occur at the edge of their geographical distribution. Moreover, Saussurea discolor is confined to the lowermost limit of its altitudinal range. Such marginality makes the area remarkable in biogeographical and conservation terms. It was revealed that a population of another rare species, Saxifraga aizoides, has possibly become extinct at this site. However, the confirmed occurrence of Saussurea discolor makes the Bili Skeli cliffs the second (in addition to Chonyi Dil Mt.) known locality of that species in the Ukrainian Carpathians.
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Claims 6
Claim · model-written summary
The cliffs host several rare calcareous saxicolous species.
- Taxon
- Arabis hornungianaArabis hornungianae0
- Outcome
- rare calcareous saxicolous speciese0
- Setting
- Bili Skelit0
Verbatim
e0“These cliffs harbour rare calcareous saxicolous species, e.g. Arabis hornungiana, Carduus defloratus subsp. glaucus, Cystopteris alpina, Epipactis atrorubens, E. helleborine, Goodyera repens, Saussurea discolor, Saxifraga luteoviridis and Thymus pulcherrimus”
t0“Bili Skeli limestone cliffs as an important hotspot of plant diversity in the Chyvchyny Mountains (Ukrainian Carpathians)”
Claim · model-written summary
Rare obligatory calcicole occurrence was associated with limestone bedrock.
- Taxon
- obligatory calcicole rare speciese0
- Outcome
- occurrence of obligatory calcicole rare speciese0
- Qualifier
- limestone bedrocke0
- Setting
- Ukrainian Carpathianse0
Not reportedDirection
Verbatim
e0“The occurrence of obligatory calcicole rare species is determined by the limestone bedrock, which is infrequent in the Ukrainian Carpathians”
t0“Bili Skeli limestone cliffs as an important hotspot of plant diversity in the Chyvchyny Mountains (Ukrainian Carpathians)”
Claim · model-written summary
Some listed species have narrow ranges and occur at distribution edges.
Verbatim
e0“Some of the listed species are narrow-range and occur at the edge of their geographical distribution”
t0“Bili Skeli limestone cliffs as an important hotspot of plant diversity in the Chyvchyny Mountains (Ukrainian Carpathians)”
Claim · model-written summary
Saussurea discolor occurs at the lowermost limit of its altitudinal range.
- Taxon
- Saussurea discolor
Saussurea discolore0
- Outcome
- confined to the lowermost limit of its altitudinal rangee0
Organisms pictured · Representative pictures of the matched species, not figures from the paper.
Verbatim
e0“Saussurea discolor is confined to the lowermost limit of its altitudinal range”
t0“Bili Skeli limestone cliffs as an important hotspot of plant diversity in the Chyvchyny Mountains (Ukrainian Carpathians)”
Claim · model-written summary
Saxifraga aizoides may have gone extinct at this site.
- Taxon
- Saxifraga aizoides
Saxifraga aizoidese0 - Outcome
- possibly become extinct at this sitee0
- Setting
- Bili Skelit0
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“a population of another rare species, Saxifraga aizoides, has possibly become extinct at this site”
t0“Bili Skeli limestone cliffs as an important hotspot of plant diversity in the Chyvchyny Mountains (Ukrainian Carpathians)”
Claim · model-written summary
Bili Skeli is the second confirmed Ukrainian Carpathian locality for Saussurea discolor.
- Taxon
- Saussurea discolor
Saussurea discolore0
- Outcome
- confirmed occurrence of Saussurea discolore0
- Value
- seconde0
- Setting
- Bili Skeli cliffse0
Organisms pictured · Representative pictures of the matched species, not figures from the paper.
Verbatim
e0“the confirmed occurrence of Saussurea discolor makes the Bili Skeli cliffs the second (in addition to Chonyi Dil Mt.) known locality of that species in the Ukrainian Carpathians”
t0“Bili Skeli limestone cliffs as an important hotspot of plant diversity in the Chyvchyny Mountains (Ukrainian Carpathians)”