Paper · 2025 · Ecological Indicators

New indicator species for associations within mesotrophic oak-hornbeam forests in Poland

Grzegorz J. Wolski; Zbigniew Celka; Jolanta Marciniuk; et al.

15
claims
22
verbatim quotes
6
citations
1
research areas
✓
Open access · cc-by

Abstract

• Research on oak-hornbeam forests reveals new pools of indicator species. • Stellario-Carpinetum is distinguished by a large group of mosses. • Other oak-hornbeam phytocenoses by high percentage of common forest species. • The current status of oak-hornbeam forests in Poland is changing. • Mosses can be highly valuable for phytosociological studies. The following oak-hornbeam plant associations have been described from Poland: Tilio - Carpinetum (TC), Galio-Carpinetum (GC) and Stellario - Carpinetum (SC). Our findings indicate that all are distinguished by a relatively high percentage of common forest species, particularly the first two. The study is the first to identify the following species as indicator species, based on SOM analysis and IndVal index: e.g. Brachythecium velutinum , Cirriphyllum piliferum , Lepidozia reptans , Mnium hornum , Plagiomnium undulatum and Brachypodium sylvaticum for SC; Ajuga reptans , Dryopteris filix-mas , Equisetum pratense , Frangula alnus , Galium odoratum , Hepatica nobilis , Luzula pilosa , Maianthemum bifolium , Milium effusum , Plagiomnium affine , Pulmonaria obscura for TC, and Asarum europaeum , Anemone ranunculoides , Corylus avellana , Galeobdolon luteum , Geum urbanum , Mercurialis perennis , Stellaria holostea and Ulmus laevis for GC. However, for Stellario-Carpinetum , our findings indicate a large pool of bryophytes that have never been considered as bioindicators. Additionally, our research indicates that while the main species composition gradient of broad-leaved forests is related to soil moisture, temperature also plays a significant role in creating a Stellario-Carpinetum – Tilio - Carpinetum gradient. The current status of oak-hornbeam forests in Poland is intriguing; our research indicates that supplementing vascular plants with bryophytes in research could change the traditional syntaxonomy of oak-hornbeam forests in Central Europe.

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Claims 15

Observationobservedoriginal resultmeasured

Claim · model-written summary

Stellario-Carpinetum is distinguished by a large group of mosses.

Taxon
mossesBryophytae4
Outcome
large group of mossese4

Verbatim

e4

“Stellario-Carpinetum is distinguished by a large group of mosses”

✓ located verbatim in the source · abstract, characters 77–141 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

All three oak-hornbeam associations have a relatively high percentage of common forest species, particularly Tilio-Carpinetum and Galio-Carpinetum.

Taxon
common forest speciese5
Outcome
relatively high percentage of common forest speciese5
Qualifier
particularly the first twoe5

Verbatim

e5

“all are distinguished by a relatively high percentage of common forest species, particularly the first two”

✓ located verbatim in the source · abstract, characters 539–645 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Brachythecium velutinum, Cirriphyllum piliferum and Lepidozia reptans are indicator species for Stellario-Carpinetum.

Taxon
Brachythecium velutinum , Cirriphyllum piliferum , Lepidozia reptansLepidozia reptansBrachytheciastrum velutinumCirriphyllum piliferume1
Outcome
indicator speciese1

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Lepidozia reptans
Lepidozia reptans · TaxonBernd Haynold · CC BY-SA 3.0 · Wikimedia Commons ↗
Brachytheciastrum velutinum
Brachytheciastrum velutinum · TaxonHermannSchachner · CC0 · Wikimedia Commons ↗
Cirriphyllum piliferum
Cirriphyllum piliferum · TaxonHermannSchachner · Public domain · Wikimedia Commons ↗

Verbatim

e1

“identify the following species as indicator species, based on SOM analysis and IndVal index: e.g. Brachythecium velutinum , Cirriphyllum piliferum , Lepidozia reptans , Mnium hornum , Plagiomnium undulatum and Brachypodium sylvaticum for SC”

✓ located verbatim in the source · abstract, characters 673–913 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Mnium hornum, Plagiomnium undulatum and Brachypodium sylvaticum are indicator species for Stellario-Carpinetum.

Taxon
Mnium hornum , Plagiomnium undulatum and Brachypodium sylvaticumBrachypodium sylvaticumMnium hornumPlagiomnium undulatume1
Outcome
indicator speciese1

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Brachypodium sylvaticum
Brachypodium sylvaticum · TaxonNo machine-readable author provided. Pere prlpz assumed (based on copyright claims). · CC BY-SA 2.5 · Wikimedia Commons ↗
Mnium hornum
Mnium hornum · TaxonHermannSchachner · CC0 · Wikimedia Commons ↗
Plagiomnium undulatum
Plagiomnium undulatum · TaxonMichael Becker · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

e1

“identify the following species as indicator species, based on SOM analysis and IndVal index: e.g. Brachythecium velutinum , Cirriphyllum piliferum , Lepidozia reptans , Mnium hornum , Plagiomnium undulatum and Brachypodium sylvaticum for SC”

✓ located verbatim in the source · abstract, characters 673–913 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Ajuga reptans, Dryopteris filix-mas and Equisetum pratense are indicator species for Tilio-Carpinetum.

Taxon
Ajuga reptans , Dryopteris filix-mas , Equisetum pratenseDryopteris filix-masAjuga reptansEquisetum pratensee2
Outcome
indicator speciese1

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Dryopteris filix-mas
Dryopteris filix-mas · TaxonB.gliwa · CC BY-SA 2.5 · Wikimedia Commons ↗
Ajuga reptans
Ajuga reptans · TaxonDidier Descouens · CC BY-SA 4.0 · Wikimedia Commons ↗
Equisetum pratense
Equisetum pratense · TaxonKristian Peters -- Fabelfroh 17:32, 12 August 2006 (UTC) · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

e1

“identify the following species as indicator species, based on SOM analysis and IndVal index: e.g. Brachythecium velutinum , Cirriphyllum piliferum , Lepidozia reptans , Mnium hornum , Plagiomnium undulatum and Brachypodium sylvaticum for SC”

✓ located verbatim in the source · abstract, characters 673–913 · show in text ↑
e2

“Ajuga reptans , Dryopteris filix-mas , Equisetum pratense , Frangula alnus , Galium odoratum , Hepatica nobilis , Luzula pilosa , Maianthemum bifolium , Milium effusum , Plagiomnium affine , Pulmonaria obscura for TC”

✓ located verbatim in the source · abstract, characters 915–1131 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Frangula alnus, Galium odoratum and Hepatica nobilis are indicator species for Tilio-Carpinetum.

Taxon
Frangula alnus , Galium odoratum , Hepatica nobilisGalium odoratumFrangula alnusHepatica nobilise2
Outcome
indicator speciese1

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Galium odoratum
Galium odoratum · TaxonJopi · Public domain · Wikimedia Commons ↗
Frangula alnus
Frangula alnus · TaxonOldMuzzle · CC BY-SA 3.0 · Wikimedia Commons ↗
Hepatica nobilis
Hepatica nobilis · TaxonKruczy89 · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

e1

“identify the following species as indicator species, based on SOM analysis and IndVal index: e.g. Brachythecium velutinum , Cirriphyllum piliferum , Lepidozia reptans , Mnium hornum , Plagiomnium undulatum and Brachypodium sylvaticum for SC”

✓ located verbatim in the source · abstract, characters 673–913 · show in text ↑
e2

“Ajuga reptans , Dryopteris filix-mas , Equisetum pratense , Frangula alnus , Galium odoratum , Hepatica nobilis , Luzula pilosa , Maianthemum bifolium , Milium effusum , Plagiomnium affine , Pulmonaria obscura for TC”

✓ located verbatim in the source · abstract, characters 915–1131 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Luzula pilosa, Maianthemum bifolium and Milium effusum are indicator species for Tilio-Carpinetum.

Taxon
Luzula pilosa , Maianthemum bifolium , Milium effusumMaianthemum bifoliumLuzula pilosaMilium effusume2
Outcome
indicator speciese1

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Maianthemum bifolium
Maianthemum bifolium · TaxonBouba at French Wikipedia · CC BY-SA 3.0 · Wikimedia Commons ↗
Luzula pilosa
Luzula pilosa · TaxonDaderot · Public domain · Wikimedia Commons ↗
Milium effusum
Milium effusum · Taxon · picture from its Wikimedia Commons categoryStefan.lefnaer · CC BY-SA 4.0 · Wikimedia Commons ↗

Verbatim

e1

“identify the following species as indicator species, based on SOM analysis and IndVal index: e.g. Brachythecium velutinum , Cirriphyllum piliferum , Lepidozia reptans , Mnium hornum , Plagiomnium undulatum and Brachypodium sylvaticum for SC”

✓ located verbatim in the source · abstract, characters 673–913 · show in text ↑
e2

“Ajuga reptans , Dryopteris filix-mas , Equisetum pratense , Frangula alnus , Galium odoratum , Hepatica nobilis , Luzula pilosa , Maianthemum bifolium , Milium effusum , Plagiomnium affine , Pulmonaria obscura for TC”

✓ located verbatim in the source · abstract, characters 915–1131 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Plagiomnium affine and Pulmonaria obscura are indicator species for Tilio-Carpinetum.

Taxon
Plagiomnium affine , Pulmonaria obscuraPlagiomnium affinePulmonaria obscurae2
Outcome
indicator speciese1

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Plagiomnium affine
Plagiomnium affine · TaxonMichael Becker · CC BY-SA 3.0 · Wikimedia Commons ↗
Pulmonaria obscura
Pulmonaria obscura · Taxon · picture from its Wikimedia Commons categoryDaniel Ballmer · CC BY-SA 4.0 · Wikimedia Commons ↗

Verbatim

e1

“identify the following species as indicator species, based on SOM analysis and IndVal index: e.g. Brachythecium velutinum , Cirriphyllum piliferum , Lepidozia reptans , Mnium hornum , Plagiomnium undulatum and Brachypodium sylvaticum for SC”

✓ located verbatim in the source · abstract, characters 673–913 · show in text ↑
e2

“Ajuga reptans , Dryopteris filix-mas , Equisetum pratense , Frangula alnus , Galium odoratum , Hepatica nobilis , Luzula pilosa , Maianthemum bifolium , Milium effusum , Plagiomnium affine , Pulmonaria obscura for TC”

✓ located verbatim in the source · abstract, characters 915–1131 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Asarum europaeum, Anemone ranunculoides and Corylus avellana are indicator species for Galio-Carpinetum.

Taxon
Asarum europaeum , Anemone ranunculoides , Corylus avellanaCorylus avellanaAsarum europaeumAnemone ranunculoidese3
Outcome
indicator speciese1

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Corylus avellana
Corylus avellana · TaxonMPF · CC BY-SA 3.0 · Wikimedia Commons ↗
Asarum europaeum
Asarum europaeum · TaxonBernd Haynold · CC BY 2.5 · Wikimedia Commons ↗
Anemone ranunculoides
Anemone ranunculoides · TaxonBogdan · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

e1

“identify the following species as indicator species, based on SOM analysis and IndVal index: e.g. Brachythecium velutinum , Cirriphyllum piliferum , Lepidozia reptans , Mnium hornum , Plagiomnium undulatum and Brachypodium sylvaticum for SC”

✓ located verbatim in the source · abstract, characters 673–913 · show in text ↑
e3

“Asarum europaeum , Anemone ranunculoides , Corylus avellana , Galeobdolon luteum , Geum urbanum , Mercurialis perennis , Stellaria holostea and Ulmus laevis for GC”

✓ located verbatim in the source · abstract, characters 1137–1300 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Galeobdolon luteum, Geum urbanum and Mercurialis perennis are indicator species for Galio-Carpinetum.

Taxon
Galeobdolon luteum , Geum urbanum , Mercurialis perennisMercurialis perennisLamium galeobdolonGeum urbanume3
Outcome
indicator speciese1

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Mercurialis perennis
Mercurialis perennis · TaxonPublic domain · Wikimedia Commons ↗
Lamium galeobdolon
Lamium galeobdolon · TaxonTeun Spaans · CC BY 2.5 · Wikimedia Commons ↗
Geum urbanum
Geum urbanum · TaxonDidier Descouens · CC BY-SA 4.0 · Wikimedia Commons ↗

Verbatim

e1

“identify the following species as indicator species, based on SOM analysis and IndVal index: e.g. Brachythecium velutinum , Cirriphyllum piliferum , Lepidozia reptans , Mnium hornum , Plagiomnium undulatum and Brachypodium sylvaticum for SC”

✓ located verbatim in the source · abstract, characters 673–913 · show in text ↑
e3

“Asarum europaeum , Anemone ranunculoides , Corylus avellana , Galeobdolon luteum , Geum urbanum , Mercurialis perennis , Stellaria holostea and Ulmus laevis for GC”

✓ located verbatim in the source · abstract, characters 1137–1300 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Stellaria holostea and Ulmus laevis are indicator species for Galio-Carpinetum.

Taxon
Stellaria holostea and Ulmus laevisUlmus laevisRabelera holosteae3
Outcome
indicator speciese1

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Ulmus laevis
Ulmus laevis · TaxonRainer Lippert · CC0 · Wikimedia Commons ↗
Rabelera holostea
Rabelera holostea · TaxonRasbak · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

e1

“identify the following species as indicator species, based on SOM analysis and IndVal index: e.g. Brachythecium velutinum , Cirriphyllum piliferum , Lepidozia reptans , Mnium hornum , Plagiomnium undulatum and Brachypodium sylvaticum for SC”

✓ located verbatim in the source · abstract, characters 673–913 · show in text ↑
e3

“Asarum europaeum , Anemone ranunculoides , Corylus avellana , Galeobdolon luteum , Geum urbanum , Mercurialis perennis , Stellaria holostea and Ulmus laevis for GC”

✓ located verbatim in the source · abstract, characters 1137–1300 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Stellario-Carpinetum has a large pool of bryophytes that have never been considered as bioindicators.

Taxon
bryophytese6
Outcome
large pool of bryophytese6

Verbatim

e6

“our findings indicate a large pool of bryophytes that have never been considered as bioindicators”

✓ located verbatim in the source · abstract, characters 1338–1435 · show in text ↑
Associationobservedoriginal resultmeasured

Claim · model-written summary

The main species composition gradient of broad-leaved forests is related to soil moisture.

Taxon
broad-leaved forestse7
Treatment
soil moisturee7
Outcome
main species composition gradiente7

Not reportedDirection

Verbatim

e7

“the main species composition gradient of broad-leaved forests is related to soil moisture, temperature also plays a significant role in creating a Stellario-Carpinetum – Tilio - Carpinetum gradient”

✓ located verbatim in the source · abstract, characters 1485–1682 · show in text ↑
Associationobservedoriginal resultsignificant (as stated)measured

Claim · model-written summary

Temperature plays a significant role in creating the Stellario-Carpinetum – Tilio-Carpinetum gradient.

Taxon
broad-leaved forestse7
Treatment
temperaturee7
Outcome
Stellario-Carpinetum – Tilio - Carpinetum gradiente7

Not reportedDirection

Verbatim

e7

“the main species composition gradient of broad-leaved forests is related to soil moisture, temperature also plays a significant role in creating a Stellario-Carpinetum – Tilio - Carpinetum gradient”

✓ located verbatim in the source · abstract, characters 1485–1682 · show in text ↑
Recommendationauthor interpretationoriginal resultbasis n/a

Claim · model-written summary

Supplementing vascular plants with bryophytes in research could change the traditional syntaxonomy of oak-hornbeam forests in Central Europe.

Taxon
vascular plantsTracheophytae0
Outcome
traditional syntaxonomy of oak-hornbeam forestse0

Verbatim

e0

“supplementing vascular plants with bryophytes in research could change the traditional syntaxonomy of oak-hornbeam forests in Central Europe”

✓ located verbatim in the source · abstract, characters 1780–1920 · show in text ↑