Paper · 2025 · Vegetation Classification and Survey

TDV-optimization: A novel numerical method for phytosociological tabulation

Tiago Monteiro-Henriques

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Open access · cc-by

Abstract

I present the Total Differential Value (TDV), an index designed for vegetation analysis based on the operational concept of differential species, as classically illustrated by Heinz Ellenberg and Dieter Mueller-Dombois. Given a phytosociological table and a grouping of its relevés, TDV is obtained by averaging the Differential Value (DiffVal) for each species in the table. DiffVal, grounded in combinatorial-discrete mathematics, quantifies the differential power of a species. The novelty of this approach lies in its distinction between two types of species absences: (i) absences from some relevés within a group and (ii) absences from all relevés representing a group. By leveraging the distribution of species absences among groups, this method effectively quantifies the differential power and distinguishes differential from non-differential species. I illustrate the computation of DiffVal and TDV and show that, because only differential species contribute to TDV, it reflects the strength of the differential species patterns in a classified table. TDV can be optimized (TDV-optimization), providing partitions of relevés. I demonstrate TDV-optimization using both an artificial and a well-known real-world data set. Key features of this method include its ability to identify patterns very closely resembling manual phytosociological tabulation and to detect reticulate patterns. TDV-optimization may lead to partitions where outlier or extreme relevés are isolated in groups; however, enforcing a minimum group size can highlight partitions with more balanced group sizes. An R package is now available, implementing DiffVal and TDV calculation as well as TDV-optimization. Abbreviations : DiffVal = Differential Value; EllPar = Ellenberg’s partition into three groups; NDR = number of discrepant relevés; NSS = number of significant species; TDV = Total Differential Value.

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Claims 8

Method / resourceauthor interpretationoriginal resultsimulated

Claim · model-written summary

DiffVal quantifies the differential power of a species.

Outcome
differential power of a speciese0

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“DiffVal, grounded in combinatorial-discrete mathematics, quantifies the differential power of a species.”

✓ located verbatim in the source · abstract, characters 376–480 · show in text ↑
Method / resourceauthor interpretationoriginal resultsimulated

Claim · model-written summary

TDV reflects the strength of differential species patterns in a classified table.

Outcome
strength of the differential species patternse0

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e0

“because only differential species contribute to TDV, it reflects the strength of the differential species patterns in a classified table”

✓ located verbatim in the source · abstract, characters 924–1060 · show in text ↑
Method / resourceobservedoriginal resultsimulated

Claim · model-written summary

TDV optimization can produce partitions of relevés.

Outcome
partitions of relevése0

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e0

“TDV can be optimized (TDV-optimization), providing partitions of relevés.”

✓ located verbatim in the source · abstract, characters 1062–1135 · show in text ↑
Method / resourceobservedoriginal resultpositivemeasured

Claim · model-written summary

The method identified patterns closely resembling manual phytosociological tabulation.

Outcome
patterns very closely resembling manual phytosociological tabulatione0
Comparator
manual phytosociological tabulatione0

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e0

“identify patterns very closely resembling manual phytosociological tabulation”

✓ located verbatim in the source · abstract, characters 1281–1358 · show in text ↑
Method / resourceauthor interpretationoriginal resultsimulated

Claim · model-written summary

TDV optimization may isolate outlier or extreme relevés into groups.

Outcome
outlier or extreme relevés are isolated in groupse0

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e0

“TDV-optimization may lead to partitions where outlier or extreme relevés are isolated in groups”

✓ located verbatim in the source · abstract, characters 1394–1489 · show in text ↑
Method / resourceauthor interpretationoriginal resultpositivesimulated

Claim · model-written summary

Enforcing a minimum group size can highlight partitions with more balanced group sizes.

Outcome
partitions with more balanced group sizese0

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e0

“enforcing a minimum group size can highlight partitions with more balanced group sizes”

✓ located verbatim in the source · abstract, characters 1500–1586 · show in text ↑
Method / resourceobservedoriginal resultbasis n/a

Claim · model-written summary

An R package implements DiffVal and TDV calculations and TDV optimization.

Outcome
R packagee0

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e0

“An R package is now available, implementing DiffVal and TDV calculation as well as TDV-optimization.”

✓ located verbatim in the source · abstract, characters 1588–1688 · show in text ↑