Paper · 2026 · International Journal of Forestry Research

Woody Plant Species Diversity in Haramaya University Main Campus, Eastern Ethiopia

Zelalem Nurlign Getahun; Anteneh Belayneh Desta; Hussen Yimer Abate

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citations
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Open access · cc-by

Abstract

Quantifying woody species diversity is essential for managing anthropogenically modified landscapes in the Ethiopian highlands. This study evaluated the floristic composition and structural diversity of woody plant species at Haramaya University, Eastern Ethiopia. Vegetation data were collected from 16 quadrats (1600 m 2 ) across two distinct managed zones: the College of Agricultural and Environmental Sciences (Site A) and the Haramaya University Model School (Site B). A total of 569 individuals representing 29 unique woody species were identified. Site B recorded a higher Shannon–Wiener Diversity Index ( H ′ = 2.78) and an Evenness Index ( J ′ = 0.928). In contrast, Site A exhibited lower diversity ( H ′ = 2.31) and an Evenness Index ( J ′ = 0.815) and a significantly higher Simpson’s Dominance Index ( D = 0.1465), which was double that of Site B ( D = 0.0752). An independent sample t ‐test performed on per‐plot diversity values confirmed that the higher structural diversity in Site B was statistically significant (t(14) = 2.45, p = 0.028). Celtis africana emerged as a major structural driver across both sites, reaching a relative density of 29.7% and a modified two‐component Importance Value Index (IVI) of 35.80% in Site A. A Jaccard Similarity Index of 27.59% indicated high beta‐diversity between the study sites, heavily influenced by the uneven introduction of exotic species. Notably, Lantana camara (IVI = 33.7%) and Jacaranda mimosifolia (IVI = 19.5%) have established substantial ecological presence, while indigenous target timber species such as Podocarpus falcatus (5.5%) and Juniperus procera (12%) exhibited lower IVIs. These findings indicate that the campus is transitioning toward a “novel ecosystem” structure. Strategic management, including the targeted suppression of invasive species and enrichment planting of native hardwoods such as Hagenia abyssinica , is recommended to maintain the historical Afromontane character of the landscape.

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Claims 16

Observationobservedoriginal resultmeasured

Claim · model-written summary

A total of 569 individuals representing 29 unique woody species were identified in 16 quadrats at Haramaya University.

Taxon
Woody Plant Speciest0
Outcome
569 individuals representing 29 unique woody species were identifiede0

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“Woody Plant Species”

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e0

“A total of 569 individuals representing 29 unique woody species were identified”

✓ located verbatim in the source · abstract, characters 475–554 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Site B (Model School) had a higher Shannon-Wiener diversity index (H′ = 2.78).

Taxon
Woody Plant Speciest0
Outcome
Shannon–Wiener Diversity Indexe0
Direction
highere0
Value
2.78e0
Setting
Site Be0

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t0

“Woody Plant Species”

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e0

“Site B recorded a higher Shannon–Wiener Diversity Index ( H ′ = 2.78) and an Evenness Index ( J ′ = 0.928)”

✓ located verbatim in the source · abstract, characters 556–662 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Site B had an evenness index of J′ = 0.928.

Taxon
Woody Plant Speciest0
Outcome
Evenness Indexe0
Value
0.928e0
Setting
Site Be0

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t0

“Woody Plant Species”

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e0

“Site B recorded a higher Shannon–Wiener Diversity Index ( H ′ = 2.78) and an Evenness Index ( J ′ = 0.928)”

✓ located verbatim in the source · abstract, characters 556–662 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Site A (College of Agricultural and Environmental Sciences) showed lower diversity (H′ = 2.31).

Taxon
Woody Plant Speciest0
Outcome
diversitye0
Direction
lowere0
Value
2.31e0
Setting
Site Ae0

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“Woody Plant Species”

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e0

“Site A exhibited lower diversity ( H ′ = 2.31) and an Evenness Index ( J ′ = 0.815)”

✓ located verbatim in the source · abstract, characters 677–760 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Site A had an evenness index of J′ = 0.815.

Taxon
Woody Plant Speciest0
Outcome
Evenness Indexe0
Value
0.815e0
Setting
Site Ae0

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“Woody Plant Species”

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e0

“Site A exhibited lower diversity ( H ′ = 2.31) and an Evenness Index ( J ′ = 0.815)”

✓ located verbatim in the source · abstract, characters 677–760 · show in text ↑
Observationobservedoriginal resultsignificant (as stated)measured

Claim · model-written summary

Site A had a significantly higher Simpson dominance index (D = 0.1465), double that of Site B (D = 0.0752).

Taxon
Woody Plant Speciest0
Outcome
Simpson’s Dominance Indexe0
Direction
highere0
Value
0.1465e0
Qualifier
0.0752e0
Comparator
Site Be0
Setting
Site Ae0

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t0

“Woody Plant Species”

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e0

“Site A exhibited lower diversity ( H ′ = 2.31) and an Evenness Index ( J ′ = 0.815) and a significantly higher Simpson’s Dominance Index ( D = 0.1465), which was double that of Site B ( D = 0.0752)”

✓ located verbatim in the source · abstract, characters 677–874 · show in text ↑
Observationobservedoriginal resultsignificant (as stated)measured

Claim · model-written summary

An independent-sample t-test on per-plot values showed the higher structural diversity at Site B was statistically significant (t(14) = 2.45, p = 0.028).

Taxon
Woody Plant Speciest0
Outcome
higher structural diversitye0
Direction
highere0
Setting
Site Be0

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t0

“Woody Plant Species”

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e0

“the higher structural diversity in Site B was statistically significant (t(14) = 2.45, p = 0.028)”

✓ located verbatim in the source · abstract, characters 960–1057 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Celtis africana was a major structural driver across both sites, with a relative density of 29.7%.

Taxon
Celtis africanaCeltis africanae0
Outcome
relative densitye0
Value
29.7%e0

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Celtis africana
Celtis africana · TaxonJMK · CC BY-SA 3.0 · Wikimedia Commons ↗

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“Woody Plant Species”

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“Celtis africana emerged as a major structural driver across both sites, reaching a relative density of 29.7%”

✓ located verbatim in the source · abstract, characters 1059–1167 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Celtis africana had a modified two-component Importance Value Index of 35.80% in Site A.

Taxon
Celtis africanaCeltis africanae1
Outcome
Importance Value Index (IVI)e0
Value
35.80%e0
Setting
Site Ae0

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Celtis africana
Celtis africana · TaxonJMK · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

t0

“Woody Plant Species”

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e1

“Celtis africana emerged as a major structural driver across both sites”

✓ located verbatim in the source · abstract, characters 1059–1129 · show in text ↑
e0

“Importance Value Index (IVI) of 35.80% in Site A”

✓ located verbatim in the source · abstract, characters 1197–1245 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

A Jaccard similarity index of 27.59% indicated high beta-diversity between the two study sites.

Taxon
Woody Plant Speciest0
Outcome
Jaccard Similarity Indexe0
Value
27.59%e0

Verbatim

t0

“Woody Plant Species”

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“A Jaccard Similarity Index of 27.59% indicated high beta-diversity between the study sites”

✓ located after whitespace/quote normalisation · abstract, characters 1247–1337 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Lantana camara (IVI = 33.7%) has established substantial ecological presence.

Taxon
Lantana camaraLantana camarae0
Outcome
substantial ecological presencee0
Value
33.7%e0

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Lantana camara
Lantana camara · TaxonW. Bulach · CC BY-SA 4.0 · Wikimedia Commons ↗

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“Woody Plant Species”

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“Notably, Lantana camara (IVI = 33.7%) and Jacaranda mimosifolia (IVI = 19.5%) have established substantial ecological presence”

✓ located verbatim in the source · abstract, characters 1404–1530 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Jacaranda mimosifolia (IVI = 19.5%) has established substantial ecological presence.

Taxon
Jacaranda mimosifoliaJacaranda mimosifoliae0
Outcome
substantial ecological presencee0
Value
19.5%e0

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Jacaranda mimosifolia
Jacaranda mimosifolia · TaxonKahuroa · Public domain · Wikimedia Commons ↗

Verbatim

t0

“Woody Plant Species”

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e0

“Notably, Lantana camara (IVI = 33.7%) and Jacaranda mimosifolia (IVI = 19.5%) have established substantial ecological presence”

✓ located verbatim in the source · abstract, characters 1404–1530 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Podocarpus falcatus, an indigenous target timber species, had a low importance value index (5.5%).

Taxon
Podocarpus falcatusAfrocarpus falcatuse0
Outcome
lower IVIse0
Direction
lowere0
Value
5.5%e0

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Afrocarpus falcatus
Afrocarpus falcatus · TaxonJMK · CC BY-SA 3.0 · Wikimedia Commons ↗

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t0

“Woody Plant Species”

✓ located verbatim in the source · title, characters 0–19 · show in text ↑
e0

“indigenous target timber species such as Podocarpus falcatus (5.5%) and Juniperus procera (12%) exhibited lower IVIs”

✓ located verbatim in the source · abstract, characters 1538–1654 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Juniperus procera, an indigenous target timber species, had a low importance value index (12%).

Taxon
Juniperus proceraJuniperus procerae0
Outcome
lower IVIse0
Direction
lowere0
Value
12%e0

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Juniperus procera
Juniperus procera · TaxonSema Tu · CC BY-SA 4.0 · Wikimedia Commons ↗

Verbatim

t0

“Woody Plant Species”

✓ located verbatim in the source · title, characters 0–19 · show in text ↑
e0

“indigenous target timber species such as Podocarpus falcatus (5.5%) and Juniperus procera (12%) exhibited lower IVIs”

✓ located verbatim in the source · abstract, characters 1538–1654 · show in text ↑
Observationauthor interpretationoriginal resultmeasured

Claim · model-written summary

The authors conclude that the campus is transitioning toward a novel ecosystem structure.

Taxon
Woody Plant Speciest0
Outcome
transitioning toward a “novel ecosystem” structuree0

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t0

“Woody Plant Species”

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“the campus is transitioning toward a “novel ecosystem” structure”

✓ located verbatim in the source · abstract, characters 1685–1749 · show in text ↑
Recommendationauthor interpretationoriginal resultbasis n/a

Claim · model-written summary

The authors recommend targeted suppression of invasive species and enrichment planting of native hardwoods to maintain the Afromontane character.

Taxon
native hardwoodse0
Outcome
Strategic managemente0

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t0

“Woody Plant Species”

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e0

“Strategic management, including the targeted suppression of invasive species and enrichment planting of native hardwoods”

✓ located verbatim in the source · abstract, characters 1751–1871 · show in text ↑