Abstract
The presence of small knobs/nodules found on the adaxial leaflet surface of some Encephalartos species was investigated in 134 cycad species of nine genera. The knobs, referred to as collicles, were found to be the result of connecting mucilage canal ‘bridges’ over vascular bundles, connecting mucilage canals situated between adjacent parallel vascular bundles. Collicles were found on some Encephalartos species only and not always on all individuals of the same species. The possible taxonomic value of these collicles is discussed.
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Claims 2
Claim · model-written summary
Collicles form from mucilage canal bridges over vascular bundles.
Verbatim
e0“The knobs, referred to as collicles, were found to be the result of connecting mucilage canal ‘bridges’ over vascular bundles, connecting mucilage canals situated between adjacent parallel vascular bundles”
t0“Cycad collicles”
Claim · model-written summary
Collicles occurred only in some Encephalartos species and varied among individuals.
- Taxon
- Encephalartos
Encephalartose0 - Outcome
- Colliclese0
- Qualifier
- not always on all individuals of the same speciese0
Not reportedDirection
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“Collicles were found on some Encephalartos species only and not always on all individuals of the same species”