Abstract
Abstract—Data on the dynamics of the ontogenetic structure of two coenopopulations (CPs) of Dracocephalum fruticulosum located in various ecological and cenotic conditions of Tuva are presented. The changes in the type of ontogenetic spectrum and the density index of individuals were analyzed. A sharp decrease in density was revealed in both CPs over 18 years. The monomodality of the ontogenetic spectrum was established during all the years of our study. There was a change in the type of the cenopopulations, namely, from transient to aging in CP 1 and from transient to mature in CP 2. The calculated demographic indicators and the rate of development reflect the unidirectional dynamics of the CP structure, namely, a successive type in CP 1 and a wave-fluctuation type in CP 2. CP analysis showed significant vulnerability of demographic indicators. Changes in the coenopopulations are due to different causes: overgrowth of communities with Selaginella sanguinolenta in CP 1, anthropogenic impact (grazing) in CP 2, and weak seed setting and germination.
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Claims 10
Claim · model-written summary
Density sharply decreased in both coenopopulations over 18 years.
- Taxon
- Dracocephalum fruticulosumDracocephalum fruticulosumt0
- Outcome
- densitye0
- Direction
- decreasee0
- Value
- 18 yearse0
Verbatim
e0“A sharp decrease in density was revealed in both CPs over 18 years.”
t0“Dracocephalum fruticulosum”
Claim · model-written summary
The ontogenetic spectrum remained monomodal in all study years.
- Taxon
- Dracocephalum fruticulosumDracocephalum fruticulosumt0
- Outcome
- monomodality of the ontogenetic spectrume0
Verbatim
e0“The monomodality of the ontogenetic spectrum was established during all the years of our study.”
t0“Dracocephalum fruticulosum”
Claim · model-written summary
CP 1 shifted from transient to aging.
- Taxon
- Dracocephalum fruticulosumDracocephalum fruticulosumt0
- Material
- CP 1e0
- Outcome
- transient to aginge0
Verbatim
e0“from transient to aging in CP 1”
t0“Dracocephalum fruticulosum”
Claim · model-written summary
CP 2 shifted from transient to mature.
- Taxon
- Dracocephalum fruticulosumDracocephalum fruticulosumt0
- Material
- CP 2e0
- Outcome
- transient to maturee0
Verbatim
e0“from transient to mature in CP 2”
t0“Dracocephalum fruticulosum”
Claim · model-written summary
CP 1 had successive structural dynamics.
- Taxon
- Dracocephalum fruticulosumDracocephalum fruticulosumt0
- Material
- CP 1e0
- Outcome
- a successive typee0
Verbatim
e0“a successive type in CP 1”
t0“Dracocephalum fruticulosum”
Claim · model-written summary
CP 2 had wave-fluctuation structural dynamics.
- Taxon
- Dracocephalum fruticulosumDracocephalum fruticulosumt0
- Material
- CP 2e0
- Outcome
- a wave-fluctuation typee0
Verbatim
e0“a wave-fluctuation type in CP 2”
t0“Dracocephalum fruticulosum”
Claim · model-written summary
Population demographic indicators were significantly vulnerable.
- Taxon
- Dracocephalum fruticulosumDracocephalum fruticulosumt0
- Outcome
- vulnerability of demographic indicatorse0
Verbatim
e0“CP analysis showed significant vulnerability of demographic indicators.”
t0“Dracocephalum fruticulosum”
Claim · model-written summary
Overgrowth by Selaginella sanguinolenta was identified as a driver of change in CP 1.
- Taxon
- Dracocephalum fruticulosumDracocephalum fruticulosumt0
- Interactor
- Selaginella sanguinolenta
Selaginella sanguinolentae0other - Outcome
- overgrowth of communitiese0
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“overgrowth of communities with Selaginella sanguinolenta in CP 1”
t0“Dracocephalum fruticulosum”
Claim · model-written summary
Grazing was identified as a driver of change in CP 2.
- Taxon
- Dracocephalum fruticulosumDracocephalum fruticulosumt0
- Outcome
- anthropogenic impacte0
- Qualifier
- grazinge0
Verbatim
e0“anthropogenic impact (grazing) in CP 2”
t0“Dracocephalum fruticulosum”
Claim · model-written summary
Weak seed setting and germination were identified as drivers of population change.
- Taxon
- Dracocephalum fruticulosumDracocephalum fruticulosumt0
- Organ / stage
- seede0
- Outcome
- weak seed setting and germinatione0
Verbatim
e0“weak seed setting and germination”
t0“Dracocephalum fruticulosum”