Paper · 2026 · Agronomy Journal

Can simple metrics predict carbon to nitrogen ratio changes in rye and triticale cover crops?

Eric B. Brennan; Richard F. Smith

10
claims
26
verbatim quotes
0
citations
3
research areas
—
Not open access

Abstract

Abstract Knowing a cover crop's carbon to nitrogen (C/N) ratio can help growers time termination to optimize N release. But growers lack simple methods to predict important C/N ratios like 20, an often‐cited “tipping point” between N mineralization and immobilization. We evaluated if Feekes growth stage, days after planting (DAP), and growing degree days (GDD) could predict C/N ratio changes of cereal cover crops. We tracked C/N changes monthly in rye ( Secale cereale L. ‘Merced’) and triticale (x Triticosecale Wittmack, ‘Pacheco’) in five biweekly trials sown from October 1 to November 30, 2021, and related these changes to farmer‐friendly metrics: DAP, GDD, and development. Growth stage and GDD were strongly correlated, as were DAP and C/N. The C/N ratios (1) increased with increasing GDD, DAP, and growth stage, (2) were similar for both cereals, and (3) became more variable over time. With exception of one trial, planted on November 15 where N was more limited, shoot biomass reached a C/N ratio of 20 when October‐planted cover crops were at Feekes 10.1 (heading, 97–103 DAP) and Feekes 8 (flag leaf just visible, 94–98 DAP) in November plantings. Excluding the 15 Nov. trial, fitted regressio curves up to ≈150 DAP showed that a C/N of 20 occurred by 94 to 102 DAP, for October‐ and November‐plantings, respectively. Accumulated GDD to reach a C/N of 20 varied with planting date ranging from approximately 650 to 900 GDD, and tended to decrease with later plantings. Our data illustrate the value and complexity of using growth stage, GDD, and DAP to predict the C/N ratio changes in cover crops. This will help farmers optimize cover crop management and comply with a new California regulation to reduce N leaching to groundwater.

Abstract from the paper’s OpenAlex record; open copy licensed public-domain. Read at the source ↗

The text the model read, with the source record’s inline markup (italics, sub- and superscripts) rendered and its character entities decoded; quote offsets refer to the raw text.

Claims 10

Associationobservedoriginal resultmeasured

Claim · model-written summary

Growth stage and growing degree days were strongly correlated in rye and triticale cover crops.

Taxon
rye and triticaleSecale cereale×Triticosecalet0
Outcome
Growth stagee0
Direction
strongly correlatede0
Comparator
GDDe0

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Secale cereale
Secale cereale · TaxonLSDSL · CC BY-SA 3.0 · Wikimedia Commons ↗
x Triticosecale
×Triticosecale · Taxon · picture from its Wikimedia Commons categoryMiquel Pujol Palol · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

t0

“rye and triticale cover crops”

✓ located verbatim in the source · title, characters 63–92 · show in text ↑
e0

“Growth stage and GDD were strongly correlated”

✓ located verbatim in the source · abstract, characters 685–730 · show in text ↑
Associationobservedoriginal resultmeasured

Claim · model-written summary

Days after planting and C/N ratio were strongly correlated in rye and triticale cover crops.

Taxon
rye and triticaleSecale cereale×Triticosecalet0
Outcome
C/Ne0
Direction
as weree0
Comparator
DAPe0

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Secale cereale
Secale cereale · TaxonLSDSL · CC BY-SA 3.0 · Wikimedia Commons ↗
x Triticosecale
×Triticosecale · Taxon · picture from its Wikimedia Commons categoryMiquel Pujol Palol · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

t0

“rye and triticale cover crops”

✓ located verbatim in the source · title, characters 63–92 · show in text ↑
e0

“as were DAP and C/N”

✓ located verbatim in the source · abstract, characters 732–751 · show in text ↑
Associationobservedoriginal resultmeasured

Claim · model-written summary

The C/N ratios of the cover crops increased with increasing GDD, DAP and growth stage.

Taxon
rye and triticaleSecale cereale×Triticosecalet0
Outcome
C/N ratiose0
Direction
increasede1
Comparator
GDD, DAP, and growth stagee1

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Secale cereale
Secale cereale · TaxonLSDSL · CC BY-SA 3.0 · Wikimedia Commons ↗
x Triticosecale
×Triticosecale · Taxon · picture from its Wikimedia Commons categoryMiquel Pujol Palol · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

t0

“rye and triticale cover crops”

✓ located verbatim in the source · title, characters 63–92 · show in text ↑
e0

“The C/N ratios”

✓ located verbatim in the source · abstract, characters 753–767 · show in text ↑
e1

“increased with increasing GDD, DAP, and growth stage”

✓ located verbatim in the source · abstract, characters 772–824 · show in text ↑
Observationobservedoriginal resultno effectmeasured

Claim · model-written summary

The C/N ratios were similar for rye and triticale.

Taxon
rye and triticaleSecale cereale×Triticosecalet0
Outcome
C/N ratiose0
Qualifier
were similar for both cerealse1

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Secale cereale
Secale cereale · TaxonLSDSL · CC BY-SA 3.0 · Wikimedia Commons ↗
x Triticosecale
×Triticosecale · Taxon · picture from its Wikimedia Commons categoryMiquel Pujol Palol · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

t0

“rye and triticale cover crops”

✓ located verbatim in the source · title, characters 63–92 · show in text ↑
e0

“The C/N ratios”

✓ located verbatim in the source · abstract, characters 753–767 · show in text ↑
e1

“were similar for both cereals”

✓ located verbatim in the source · abstract, characters 830–859 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

The C/N ratios became more variable over time.

Taxon
rye and triticaleSecale cereale×Triticosecalet0
Outcome
C/N ratiose0
Qualifier
became more variable over timee1

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Secale cereale
Secale cereale · TaxonLSDSL · CC BY-SA 3.0 · Wikimedia Commons ↗
x Triticosecale
×Triticosecale · Taxon · picture from its Wikimedia Commons categoryMiquel Pujol Palol · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

t0

“rye and triticale cover crops”

✓ located verbatim in the source · title, characters 63–92 · show in text ↑
e0

“The C/N ratios”

✓ located verbatim in the source · abstract, characters 753–767 · show in text ↑
e1

“became more variable over time”

✓ located verbatim in the source · abstract, characters 869–899 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Shoot biomass of October-planted cover crops reached a C/N ratio of 20 at Feekes 10.1 (heading), 97-103 days after planting.

Taxon
rye and triticaleSecale cereale×Triticosecalet0
Outcome
C/N ratio of 20e0
Value
97–103 DAPe1
Qualifier
October‐planted cover crops were at Feekes 10.1e1

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Secale cereale
Secale cereale · TaxonLSDSL · CC BY-SA 3.0 · Wikimedia Commons ↗
x Triticosecale
×Triticosecale · Taxon · picture from its Wikimedia Commons categoryMiquel Pujol Palol · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

t0

“rye and triticale cover crops”

✓ located verbatim in the source · title, characters 63–92 · show in text ↑
e0

“shoot biomass reached a C/N ratio of 20”

✓ located verbatim in the source · abstract, characters 979–1018 · show in text ↑
e1

“October‐planted cover crops were at Feekes 10.1 (heading, 97–103 DAP)”

✓ located verbatim in the source · abstract, characters 1024–1093 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Shoot biomass of November-planted cover crops reached a C/N ratio of 20 at Feekes 8 (flag leaf just visible), 94-98 days after planting.

Taxon
rye and triticaleSecale cereale×Triticosecalet0
Outcome
C/N ratio of 20e0
Value
94–98 DAPe1
Qualifier
Feekes 8 (flag leaf just visible, 94–98 DAP) in November plantingse1

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Secale cereale
Secale cereale · TaxonLSDSL · CC BY-SA 3.0 · Wikimedia Commons ↗
x Triticosecale
×Triticosecale · Taxon · picture from its Wikimedia Commons categoryMiquel Pujol Palol · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

t0

“rye and triticale cover crops”

✓ located verbatim in the source · title, characters 63–92 · show in text ↑
e0

“shoot biomass reached a C/N ratio of 20”

✓ located verbatim in the source · abstract, characters 979–1018 · show in text ↑
e1

“Feekes 8 (flag leaf just visible, 94–98 DAP) in November plantings”

✓ located verbatim in the source · abstract, characters 1098–1164 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Fitted regression curves showed a C/N of 20 occurring by 94 to 102 days after planting for October and November plantings, respectively.

Taxon
rye and triticaleSecale cereale×Triticosecalet0
Outcome
C/N of 20e0
Value
94 to 102 DAPe0
Qualifier
for October‐ and November‐plantingse0

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Secale cereale
Secale cereale · TaxonLSDSL · CC BY-SA 3.0 · Wikimedia Commons ↗
x Triticosecale
×Triticosecale · Taxon · picture from its Wikimedia Commons categoryMiquel Pujol Palol · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

t0

“rye and triticale cover crops”

✓ located verbatim in the source · title, characters 63–92 · show in text ↑
e0

“a C/N of 20 occurred by 94 to 102 DAP, for October‐ and November‐plantings, respectively”

✓ located verbatim in the source · abstract, characters 1246–1334 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Accumulated GDD to reach a C/N of 20 varied with planting date, ranging from approximately 650 to 900 GDD.

Taxon
rye and triticaleSecale cereale×Triticosecalet0
Outcome
Accumulated GDD to reach a C/N of 20e0
Value
approximately 650 to 900 GDDe0
Qualifier
varied with planting datee0

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Secale cereale
Secale cereale · TaxonLSDSL · CC BY-SA 3.0 · Wikimedia Commons ↗
x Triticosecale
×Triticosecale · Taxon · picture from its Wikimedia Commons categoryMiquel Pujol Palol · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

t0

“rye and triticale cover crops”

✓ located verbatim in the source · title, characters 63–92 · show in text ↑
e0

“Accumulated GDD to reach a C/N of 20 varied with planting date ranging from approximately 650 to 900 GDD”

✓ located verbatim in the source · abstract, characters 1336–1440 · show in text ↑
Intervention effectobservedoriginal resultmeasured

Claim · model-written summary

Accumulated GDD to reach a C/N of 20 tended to decrease with later plantings.

Taxon
rye and triticaleSecale cereale×Triticosecalet0
Treatment
later plantingse1
Outcome
Accumulated GDD to reach a C/N of 20e0
Direction
decreasee1

Not reportedComparator

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Secale cereale
Secale cereale · TaxonLSDSL · CC BY-SA 3.0 · Wikimedia Commons ↗
x Triticosecale
×Triticosecale · Taxon · picture from its Wikimedia Commons categoryMiquel Pujol Palol · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

t0

“rye and triticale cover crops”

✓ located verbatim in the source · title, characters 63–92 · show in text ↑
e0

“Accumulated GDD to reach a C/N of 20”

✓ located verbatim in the source · abstract, characters 1336–1372 · show in text ↑
e1

“tended to decrease with later plantings”

✓ located verbatim in the source · abstract, characters 1446–1485 · show in text ↑