Paper · 2025 · Horticulturae

Impacts of Tree Thinning on Overall Productivity in Densely Planted Walnut Orchards

Qian Ye; Qinyang Yue; Yingxia Zhang; et al.

20
claims
71
verbatim quotes
2
citations
4
research areas
✓
Open access · cc-by

Abstract

To effectively address the issues of poor ventilation, light deficiency, increased pest and disease pressure, and declining fruit quality in closed-canopy walnut orchards, this study was conducted in a standard, densely planted ‘Xinwen 185’ walnut orchard. Three treatments were established: an unthinned control (CK), a 1-year thinning treatment (T1), and a 2-year thinning treatment (T2). All parameters were uniformly investigated during the 2023 growing season to analyze the effects of thinning on orchard population structure, microenvironment, leaf physiological characteristics, fruit quality, and yield. The results demonstrated that tree thinning significantly optimized the population structure: crown width expanded by 6.22–6.76 m, light transmittance increased to 27.74–33.64%, and orchard coverage decreased from 100% to 75.94–80.51%. The microenvironment was improved: inter-row temperature increased by 2.34–4.08 °C, light intensity increased by 5.38–25.29%, and relative humidity decreased by 2.15–3.30%. Furthermore, leaf physiological functions were activated: in the T2 treatment, the chlorophyll content in outer-canopy leaves increased by 15.23% and 12.45% at the kernel-hardening and maturity stages, respectively; the leaf carbon-to-nitrogen ratio increased by 18.67%; the net photosynthetic rate (Pn) during fruit expansion increased by 34.21–46.10%; and the intercellular CO2 concentration (Ci) decreased by 10.18–10.31%. Fruit quality and yield were synergistically enhanced: single fruit weight increased by 23.39~37.94%, and kernel weight increased by 26.79–41.13%. The total sugar content in inner-canopy fruits increased by 16.50–16.67%, while the protein and fat content in outer-canopy fruits increased by 0.69–12.50% and 0.60–2.18%, respectively. Yield exhibited a “short-term adjustment and long-term gain” pattern: the T2 treatment (after 2 years of thinning) achieved a yield of 5.26 t·ha−1, which was 20.38% higher than the CK. The rates of diseased fruit and empty shells decreased by 65.71% and 93.22%, respectively, and the premium fruit rate reached 90.60%. This study confirms that tree thinning is an effective measure for improving the growing environment and enhancing overall productivity in closed-canopy walnut orchards, providing a scientific basis for sustainable orchard management and increased orchard profitability.

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Claims 20

Intervention effectobservedoriginal resultpositivemeasured

Claim · model-written summary

Tree thinning expanded crown width by 6.22-6.76 m.

Taxon
Walnutt1
Treatment
Tree Thinningt0
Outcome
crown widthe0
Direction
expandede0
Value
6.22–6.76 me0
Comparator
an unthinned control (CK)e1

Verbatim

t1

“Walnut Orchards”

✓ located verbatim in the source · title, characters 68–83 · show in text ↑
e0

“crown width expanded by 6.22–6.76 m”

✓ located verbatim in the source · abstract, characters 707–742 · show in text ↑
t0

“Tree Thinning”

✓ located verbatim in the source · title, characters 11–24 · show in text ↑
e1

“an unthinned control (CK)”

✓ located verbatim in the source · abstract, characters 292–317 · show in text ↑
Intervention effectobservedoriginal resultpositivemeasured

Claim · model-written summary

Tree thinning increased light transmittance to 27.74-33.64%.

Taxon
Walnutt1
Treatment
Tree Thinningt0
Outcome
light transmittancee0
Direction
increasede0
Value
27.74–33.64%e0
Comparator
an unthinned control (CK)e1

Verbatim

t1

“Walnut Orchards”

✓ located verbatim in the source · title, characters 68–83 · show in text ↑
e0

“light transmittance increased to 27.74–33.64%”

✓ located verbatim in the source · abstract, characters 744–789 · show in text ↑
t0

“Tree Thinning”

✓ located verbatim in the source · title, characters 11–24 · show in text ↑
e1

“an unthinned control (CK)”

✓ located verbatim in the source · abstract, characters 292–317 · show in text ↑
Intervention effectobservedoriginal resultmeasured

Claim · model-written summary

Tree thinning reduced orchard coverage from 100% to 75.94-80.51%.

Taxon
Walnutt1
Treatment
Tree Thinningt0
Outcome
orchard coveragee0
Direction
decreasede0
Value
75.94–80.51%e0
Comparator
an unthinned control (CK)e1

Verbatim

t1

“Walnut Orchards”

✓ located verbatim in the source · title, characters 68–83 · show in text ↑
e0

“orchard coverage decreased from 100% to 75.94–80.51%”

✓ located verbatim in the source · abstract, characters 795–847 · show in text ↑
t0

“Tree Thinning”

✓ located verbatim in the source · title, characters 11–24 · show in text ↑
e1

“an unthinned control (CK)”

✓ located verbatim in the source · abstract, characters 292–317 · show in text ↑
Intervention effectobservedoriginal resultmeasured

Claim · model-written summary

Tree thinning raised inter-row temperature by 2.34-4.08 degrees C.

Taxon
Walnutt1
Treatment
Tree Thinningt0
Outcome
inter-row temperaturee0
Direction
increasede0
Value
2.34–4.08 °Ce0
Comparator
an unthinned control (CK)e1

Verbatim

t1

“Walnut Orchards”

✓ located verbatim in the source · title, characters 68–83 · show in text ↑
e0

“inter-row temperature increased by 2.34–4.08 °C”

✓ located verbatim in the source · abstract, characters 884–931 · show in text ↑
t0

“Tree Thinning”

✓ located verbatim in the source · title, characters 11–24 · show in text ↑
e1

“an unthinned control (CK)”

✓ located verbatim in the source · abstract, characters 292–317 · show in text ↑
Intervention effectobservedoriginal resultpositivemeasured

Claim · model-written summary

Tree thinning raised light intensity by 5.38-25.29%.

Taxon
Walnutt1
Treatment
Tree Thinningt0
Outcome
light intensitye0
Direction
increasede0
Value
5.38–25.29%e0
Comparator
an unthinned control (CK)e1

Verbatim

t1

“Walnut Orchards”

✓ located verbatim in the source · title, characters 68–83 · show in text ↑
e0

“light intensity increased by 5.38–25.29%”

✓ located verbatim in the source · abstract, characters 933–973 · show in text ↑
t0

“Tree Thinning”

✓ located verbatim in the source · title, characters 11–24 · show in text ↑
e1

“an unthinned control (CK)”

✓ located verbatim in the source · abstract, characters 292–317 · show in text ↑
Intervention effectobservedoriginal resultpositivemeasured

Claim · model-written summary

Tree thinning lowered relative humidity by 2.15-3.30%.

Taxon
Walnutt1
Treatment
Tree Thinningt0
Outcome
relative humiditye0
Direction
decreasede0
Value
2.15–3.30%e0
Comparator
an unthinned control (CK)e1

Verbatim

t1

“Walnut Orchards”

✓ located verbatim in the source · title, characters 68–83 · show in text ↑
e0

“relative humidity decreased by 2.15–3.30%”

✓ located verbatim in the source · abstract, characters 979–1020 · show in text ↑
t0

“Tree Thinning”

✓ located verbatim in the source · title, characters 11–24 · show in text ↑
e1

“an unthinned control (CK)”

✓ located verbatim in the source · abstract, characters 292–317 · show in text ↑
Intervention effectobservedoriginal resultpositivemeasured

Claim · model-written summary

In the T2 treatment, chlorophyll content in outer-canopy leaves increased by 15.23% at the kernel-hardening stage.

Taxon
Walnutt1
Organ / stage
outer-canopy leavese0
Treatment
T2 treatmente0
Outcome
chlorophyll contente0
Direction
increasede0
Value
15.23%e0
Qualifier
kernel-hardeninge0
Comparator
an unthinned control (CK)e1

Verbatim

t1

“Walnut Orchards”

✓ located verbatim in the source · title, characters 68–83 · show in text ↑
e0

“in the T2 treatment, the chlorophyll content in outer-canopy leaves increased by 15.23% and 12.45% at the kernel-hardening and maturity stages, respectively”

✓ located verbatim in the source · abstract, characters 1080–1236 · show in text ↑
e1

“an unthinned control (CK)”

✓ located verbatim in the source · abstract, characters 292–317 · show in text ↑
Intervention effectobservedoriginal resultpositivemeasured

Claim · model-written summary

In the T2 treatment, chlorophyll content in outer-canopy leaves increased by 12.45% at the maturity stage.

Taxon
Walnutt1
Organ / stage
outer-canopy leavese0
Treatment
T2 treatmente0
Outcome
chlorophyll contente0
Direction
increasede0
Value
12.45%e0
Qualifier
maturitye0
Comparator
an unthinned control (CK)e1

Verbatim

t1

“Walnut Orchards”

✓ located verbatim in the source · title, characters 68–83 · show in text ↑
e0

“in the T2 treatment, the chlorophyll content in outer-canopy leaves increased by 15.23% and 12.45% at the kernel-hardening and maturity stages, respectively”

✓ located verbatim in the source · abstract, characters 1080–1236 · show in text ↑
e1

“an unthinned control (CK)”

✓ located verbatim in the source · abstract, characters 292–317 · show in text ↑
Intervention effectobservedoriginal resultmeasured

Claim · model-written summary

Thinning increased the leaf carbon-to-nitrogen ratio by 18.67%.

Taxon
Walnutt1
Treatment
Tree Thinningt0
Outcome
leaf carbon-to-nitrogen ratioe0
Direction
increasede0
Value
18.67%e0
Comparator
an unthinned control (CK)e1

Verbatim

t1

“Walnut Orchards”

✓ located verbatim in the source · title, characters 68–83 · show in text ↑
e0

“the leaf carbon-to-nitrogen ratio increased by 18.67%”

✓ located verbatim in the source · abstract, characters 1238–1291 · show in text ↑
t0

“Tree Thinning”

✓ located verbatim in the source · title, characters 11–24 · show in text ↑
e1

“an unthinned control (CK)”

✓ located verbatim in the source · abstract, characters 292–317 · show in text ↑
Intervention effectobservedoriginal resultpositivemeasured

Claim · model-written summary

Thinning increased net photosynthetic rate during fruit expansion by 34.21-46.10%.

Taxon
Walnutt1
Treatment
Tree Thinningt0
Outcome
net photosynthetic rate (Pn)e0
Direction
increasede0
Value
34.21–46.10%e0
Comparator
an unthinned control (CK)e1

Verbatim

t1

“Walnut Orchards”

✓ located verbatim in the source · title, characters 68–83 · show in text ↑
e0

“the net photosynthetic rate (Pn) during fruit expansion increased by 34.21–46.10%”

✓ located verbatim in the source · abstract, characters 1293–1374 · show in text ↑
t0

“Tree Thinning”

✓ located verbatim in the source · title, characters 11–24 · show in text ↑
e1

“an unthinned control (CK)”

✓ located verbatim in the source · abstract, characters 292–317 · show in text ↑
Intervention effectobservedoriginal resultmeasured

Claim · model-written summary

Thinning decreased intercellular CO2 concentration by 10.18-10.31%.

Taxon
Walnutt1
Treatment
Tree Thinningt0
Outcome
intercellular CO2 concentration (Ci)e0
Direction
decreasede0
Value
10.18–10.31%e0
Comparator
an unthinned control (CK)e1

Verbatim

t1

“Walnut Orchards”

✓ located verbatim in the source · title, characters 68–83 · show in text ↑
e0

“the intercellular CO2 concentration (Ci) decreased by 10.18–10.31%”

✓ located verbatim in the source · abstract, characters 1380–1446 · show in text ↑
t0

“Tree Thinning”

✓ located verbatim in the source · title, characters 11–24 · show in text ↑
e1

“an unthinned control (CK)”

✓ located verbatim in the source · abstract, characters 292–317 · show in text ↑
Intervention effectobservedoriginal resultpositivemeasured

Claim · model-written summary

Thinning increased single fruit weight by 23.39-37.94%.

Taxon
Walnutt1
Treatment
Tree Thinningt0
Outcome
single fruit weighte0
Direction
increasede0
Value
23.39~37.94%e0
Comparator
an unthinned control (CK)e1

Verbatim

t1

“Walnut Orchards”

✓ located verbatim in the source · title, characters 68–83 · show in text ↑
e0

“single fruit weight increased by 23.39~37.94%”

✓ located verbatim in the source · abstract, characters 1503–1548 · show in text ↑
t0

“Tree Thinning”

✓ located verbatim in the source · title, characters 11–24 · show in text ↑
e1

“an unthinned control (CK)”

✓ located verbatim in the source · abstract, characters 292–317 · show in text ↑
Intervention effectobservedoriginal resultpositivemeasured

Claim · model-written summary

Thinning increased kernel weight by 26.79-41.13%.

Taxon
Walnutt1
Treatment
Tree Thinningt0
Outcome
kernel weighte0
Direction
increasede0
Value
26.79–41.13%e0
Comparator
an unthinned control (CK)e1

Verbatim

t1

“Walnut Orchards”

✓ located verbatim in the source · title, characters 68–83 · show in text ↑
e0

“kernel weight increased by 26.79–41.13%”

✓ located verbatim in the source · abstract, characters 1554–1593 · show in text ↑
t0

“Tree Thinning”

✓ located verbatim in the source · title, characters 11–24 · show in text ↑
e1

“an unthinned control (CK)”

✓ located verbatim in the source · abstract, characters 292–317 · show in text ↑
Intervention effectobservedoriginal resultpositivemeasured

Claim · model-written summary

Thinning increased total sugar content in inner-canopy fruits by 16.50-16.67%.

Taxon
Walnutt1
Organ / stage
inner-canopy fruitse0
Treatment
Tree Thinningt0
Outcome
total sugar contente0
Direction
increasede0
Value
16.50–16.67%e0
Comparator
an unthinned control (CK)e1

Verbatim

t1

“Walnut Orchards”

✓ located verbatim in the source · title, characters 68–83 · show in text ↑
e0

“The total sugar content in inner-canopy fruits increased by 16.50–16.67%”

✓ located verbatim in the source · abstract, characters 1595–1667 · show in text ↑
t0

“Tree Thinning”

✓ located verbatim in the source · title, characters 11–24 · show in text ↑
e1

“an unthinned control (CK)”

✓ located verbatim in the source · abstract, characters 292–317 · show in text ↑
Intervention effectobservedoriginal resultpositivemeasured

Claim · model-written summary

Thinning increased protein and fat content in outer-canopy fruits by 0.69-12.50% and 0.60-2.18%, respectively.

Taxon
Walnutt1
Organ / stage
outer-canopy fruitse0
Treatment
Tree Thinningt0
Outcome
protein and fat contente0
Direction
increasede0
Value
0.69–12.50% and 0.60–2.18%e0
Comparator
an unthinned control (CK)e1

Verbatim

t1

“Walnut Orchards”

✓ located verbatim in the source · title, characters 68–83 · show in text ↑
e0

“the protein and fat content in outer-canopy fruits increased by 0.69–12.50% and 0.60–2.18%, respectively”

✓ located verbatim in the source · abstract, characters 1675–1779 · show in text ↑
t0

“Tree Thinning”

✓ located verbatim in the source · title, characters 11–24 · show in text ↑
e1

“an unthinned control (CK)”

✓ located verbatim in the source · abstract, characters 292–317 · show in text ↑
Intervention effectobservedoriginal resultpositivemeasured

Claim · model-written summary

After 2 years of thinning (T2) yield reached 5.26 t/ha, 20.38% higher than the control.

Taxon
Walnutt1
Treatment
T2 treatmente0
Outcome
yielde0
Direction
highere0
Value
5.26 t·ha−1e0
Qualifier
20.38%e0
Comparator
the CKe0

Verbatim

t1

“Walnut Orchards”

✓ located verbatim in the source · title, characters 68–83 · show in text ↑
e0

“the T2 treatment (after 2 years of thinning) achieved a yield of 5.26 t·ha−1, which was 20.38% higher than the CK”

✓ located verbatim in the source · abstract, characters 1851–1964 · show in text ↑
Observationobservedoriginal resultmeasured

Claim · model-written summary

Yield followed a short-term adjustment and long-term gain pattern under thinning.

Taxon
Walnutt1
Outcome
Yielde0
Qualifier
short-term adjustment and long-term gaine0

Verbatim

t1

“Walnut Orchards”

✓ located verbatim in the source · title, characters 68–83 · show in text ↑
e0

“Yield exhibited a “short-term adjustment and long-term gain” pattern”

✓ located verbatim in the source · abstract, characters 1781–1849 · show in text ↑
Intervention effectobservedoriginal resultpositivemeasured

Claim · model-written summary

Thinning lowered the rates of diseased fruit and empty shells by 65.71% and 93.22%, respectively.

Taxon
Walnutt1
Treatment
Tree Thinningt0
Outcome
rates of diseased fruit and empty shellse0
Direction
decreasede0
Value
65.71% and 93.22%e0
Comparator
an unthinned control (CK)e1

Verbatim

t1

“Walnut Orchards”

✓ located verbatim in the source · title, characters 68–83 · show in text ↑
e0

“The rates of diseased fruit and empty shells decreased by 65.71% and 93.22%, respectively”

✓ located verbatim in the source · abstract, characters 1966–2055 · show in text ↑
t0

“Tree Thinning”

✓ located verbatim in the source · title, characters 11–24 · show in text ↑
e1

“an unthinned control (CK)”

✓ located verbatim in the source · abstract, characters 292–317 · show in text ↑
Intervention effectobservedoriginal resultpositivemeasured

Claim · model-written summary

Under thinning the premium fruit rate reached 90.60%.

Taxon
Walnutt1
Treatment
Tree Thinningt0
Outcome
premium fruit ratee0
Value
90.60%e0
Comparator
an unthinned control (CK)e1

Not reportedDirection

Verbatim

t1

“Walnut Orchards”

✓ located verbatim in the source · title, characters 68–83 · show in text ↑
e0

“the premium fruit rate reached 90.60%”

✓ located verbatim in the source · abstract, characters 2061–2098 · show in text ↑
t0

“Tree Thinning”

✓ located verbatim in the source · title, characters 11–24 · show in text ↑
e1

“an unthinned control (CK)”

✓ located verbatim in the source · abstract, characters 292–317 · show in text ↑
Recommendationauthor interpretationoriginal resultpositivebasis n/a

Claim · model-written summary

The authors conclude that tree thinning is an effective measure for improving the growing environment and productivity of closed-canopy walnut orchards.

Taxon
walnute0
Outcome
improving the growing environment and enhancing overall productivitye0

Verbatim

e0

“tree thinning is an effective measure for improving the growing environment and enhancing overall productivity in closed-canopy walnut orchards”

✓ located verbatim in the source · abstract, characters 2125–2268 · show in text ↑

pruning · physiology: photosynthesis · Juglans (walnuts)Results agree

2 papers: 2 positive, 0 negative or no effect

1 paper in the same direction as this paper

  1. positive
See all 2 papers →

pruning · quality: protein · Juglans (walnuts)Results agree

2 papers: 2 positive, 0 negative or no effect

1 paper in the same direction as this paper

  1. positive
See all 2 papers →

pruning · yield · Juglans (walnuts)Results agree

5 papers: 5 positive, 0 negative or no effect

4 papers in the same direction as this paper

  1. positive
  2. positive
  3. positive

1 paper more in this group

See all 5 papers →

Claims are grouped by organism, treatment and what was measured or targeted. A paper counts as positive when any of its claims in the group is positive.