Abstract
Diverse cultivars of winter wheat ( Triticum aestivum L.) were grown in the field in 1993/94 and 1994/95 at Reading UK in temperature gradient tunnels at normal atmospheric ( c . 370) or elevated CO 2 concentration ( c . 700 μmol CO 2 mol −1 air). In 1993/94, grain yield of cv. Avalon was insensitive to mean temperature (between 8·8 and 10·9°C), while elevated CO 2 increased yield by 1·3 t ha −1 (12·6%). In all other cultivars, warming reduced grain yield and CO 2 increased grain yield. In 1993/94, in cvs Galahad and Mercia the effects of CO 2 and temperature on yield were additive. However, for cv. Hereward in both years and for cv. Soissons in 1994/95, there were negative interactions between the effects of CO 2 and temperature on yield: the maximum benefit of doubling CO 2 to grain yield, 4·5 and 2·7 t ha −1 (65 and 29%) respectively, occurred at cooler temperatures; there was no benefit from doubling CO 2 (i.e. 0%) once the temperature had increased above the seasonal mean by 2·2–2·6°C in cv. Hereward and by 1·3°C in cv. Soissons. The beneficial effect of doubling CO 2 on grain yield in cvs Galahad, Hereward, Mercia and Soissons was negated by an increase in mean seasonal temperature of only 0·7–2·0°C. Warming decreased root dry mass at anthesis in 1994/95 while it increased at elevated CO 2 (49 and 186%, coolest and warmest regime, respectively). Carbon partitioned to roots declined progressively with warming, while at elevated CO 2 there was an average of 56% increase in allocation to roots. The relative impacts of both CO 2 and temperature were greater on root dry mass than on either grain yield or total above-ground biomass, while the effects on grain and biomass yield varied considerably between cultivars, suggesting that the impact of rising CO 2 and temperature are likely to be dependent on cultivar.
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Claims 17
Claim · model-written summary
Avalon grain yield was insensitive to mean temperature from 8·8 to 10·9°C.
- Taxon
- winter wheat
Triticum aestivumt0 - Material
- cv. Avalone0
- Treatment
- mean temperaturee0
- Outcome
- grain yielde0
- Direction
- insensitivee0
- Qualifier
- between 8·8 and 10·9°Ce0
Not reportedComparator
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“grain yield of cv. Avalon was insensitive to mean temperature (between 8·8 and 10·9°C)”
t0“winter wheat”
Claim · model-written summary
Elevated CO2 increased Avalon grain yield by 1·3 t ha −1.
- Taxon
- winter wheat
Triticum aestivumt0 - Treatment
- elevated CO 2e0
- Outcome
- yielde0
- Direction
- increasede0
- Value
- 1·3 t ha −1e0
Not reportedMaterialComparator
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“elevated CO 2 increased yield by 1·3 t ha −1 (12·6%)”
t0“winter wheat”
Claim · model-written summary
Warming reduced grain yield in the other cultivars.
- Taxon
- winter wheat
Triticum aestivumt0 - Treatment
- warminge0
- Outcome
- grain yielde0
- Direction
- reducede0
- Qualifier
- all other cultivarse0
Not reportedComparator
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“In all other cultivars, warming reduced grain yield”
t0“winter wheat”
Claim · model-written summary
Elevated CO2 increased grain yield in the other cultivars.
- Taxon
- winter wheat
Triticum aestivumt0 - Treatment
- CO 2e0
- Outcome
- grain yielde0
- Direction
- increasede0
Not reportedComparator
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“CO 2 increased grain yield”
t0“winter wheat”
Claim · model-written summary
CO2 and temperature effects on Galahad and Mercia yield were additive.
- Taxon
- winter wheat
Triticum aestivumt0 - Material
- cvs Galahad and Merciae0
- Outcome
- effects of CO 2 and temperature on yield were additivee0
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“in cvs Galahad and Mercia the effects of CO 2 and temperature on yield were additive”
t0“winter wheat”
Claim · model-written summary
Hereward and Soissons showed negative CO2 by temperature interactions for yield.
- Taxon
- winter wheat
Triticum aestivumt0 - Material
- cv. Herewarde0
- Outcome
- negative interactions between the effects of CO 2 and temperature on yielde0
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“for cv. Hereward in both years and for cv. Soissons in 1994/95, there were negative interactions between the effects of CO 2 and temperature on yield”
t0“winter wheat”
Claim · model-written summary
The maximum CO2 benefit to Hereward grain yield was 4·5 t ha −1 at cooler temperatures.
- Taxon
- winter wheat
Triticum aestivumt0 - Treatment
- doubling CO 2e0
- Outcome
- grain yielde0
- Direction
- maximume0
- Value
- 4·5e0
- Qualifier
- cooler temperaturese0
Not reportedComparator
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“maximum benefit of doubling CO 2 to grain yield, 4·5 and 2·7 t ha −1 (65 and 29%) respectively, occurred at cooler temperatures”
t0“winter wheat”
Claim · model-written summary
The maximum CO2 benefit to Soissons grain yield was 2·7 t ha −1 at cooler temperatures.
- Taxon
- winter wheat
Triticum aestivumt0 - Treatment
- doubling CO 2e0
- Outcome
- grain yielde0
- Direction
- maximume0
- Value
- 2·7 t ha −1e0
- Qualifier
- cooler temperaturese0
Not reportedComparator
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“maximum benefit of doubling CO 2 to grain yield, 4·5 and 2·7 t ha −1 (65 and 29%) respectively, occurred at cooler temperatures”
t0“winter wheat”
Claim · model-written summary
Doubling CO2 brought no yield benefit for Hereward above a 2·2–2·6°C temperature increase.
- Taxon
- winter wheat
Triticum aestivumt0 - Material
- cv. Herewarde0
- Treatment
- doubling CO 2e0
- Outcome
- benefite0
- Direction
- no benefite0
- Value
- 0%e0
- Qualifier
- 2·2–2·6°Ce0
Not reportedComparator
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“there was no benefit from doubling CO 2 (i.e. 0%) once the temperature had increased above the seasonal mean by 2·2–2·6°C in cv. Hereward”
t0“winter wheat”
Claim · model-written summary
Doubling CO2 brought no yield benefit for Soissons above a 1·3°C increase.
- Taxon
- winter wheat
Triticum aestivumt0 - Material
- cv. Soissonse0
- Treatment
- doubling CO 2e0
- Outcome
- benefite0
- Direction
- no benefite0
- Value
- 0%e0
- Qualifier
- 1·3°Ce0
Not reportedComparator
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“there was no benefit from doubling CO 2 (i.e. 0%) once the temperature had increased above the seasonal mean by 2·2–2·6°C in cv. Hereward and by 1·3°C in cv. Soissons”
t0“winter wheat”
Claim · model-written summary
A 0·7–2·0°C warming negated the CO2 yield benefit in four cultivars.
- Taxon
- winter wheat
Triticum aestivumt0 - Treatment
- increase in mean seasonal temperaturee0
- Outcome
- beneficial effect of doubling CO 2 on grain yielde0
- Direction
- negatede0
- Value
- 0·7–2·0°Ce0
Not reportedComparator
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“beneficial effect of doubling CO 2 on grain yield in cvs Galahad, Hereward, Mercia and Soissons was negated by an increase in mean seasonal temperature of only 0·7–2·0°C”
t0“winter wheat”
Claim · model-written summary
Warming decreased root dry mass at anthesis.
- Taxon
- winter wheat
Triticum aestivumt0 - Organ / stage
- anthesise0
- Treatment
- Warminge0
- Outcome
- root dry masse0
- Direction
- decreasede0
Not reportedComparator
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“Warming decreased root dry mass at anthesis in 1994/95”
t0“winter wheat”
Claim · model-written summary
Elevated CO2 increased root dry mass.
- Taxon
- winter wheat
Triticum aestivumt0 - Treatment
- elevated CO 2e0
- Outcome
- root dry masse0
- Direction
- increasede0
- Value
- 49 and 186%e0
Not reportedComparator
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“Warming decreased root dry mass at anthesis in 1994/95 while it increased at elevated CO 2 (49 and 186%, coolest and warmest regime, respectively)”
t0“winter wheat”
Claim · model-written summary
Warming progressively reduced carbon allocation to roots.
- Taxon
- winter wheat
Triticum aestivumt0 - Treatment
- warminge0
- Outcome
- Carbon partitioned to rootse0
- Direction
- declined progressivelye0
Not reportedComparator
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“Carbon partitioned to roots declined progressively with warming”
t0“winter wheat”
Claim · model-written summary
Elevated CO2 increased root carbon allocation by 56% on average.
- Taxon
- winter wheat
Triticum aestivumt0 - Treatment
- elevated CO 2e0
- Outcome
- allocation to rootse0
- Direction
- increasee0
- Value
- 56%e0
Not reportedComparator
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“at elevated CO 2 there was an average of 56% increase in allocation to roots”
t0“winter wheat”
Claim · model-written summary
Root dry mass responded more strongly than grain yield or above-ground biomass to CO2 and temperature.
- Taxon
- winter wheat
Triticum aestivumt0 - Outcome
- root dry masse0
- Direction
- greatere0
- Comparator
- grain yield or total above-ground biomasse0
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“relative impacts of both CO 2 and temperature were greater on root dry mass than on either grain yield or total above-ground biomass”
t0“winter wheat”
Claim · model-written summary
Effects on grain and biomass yield varied considerably among cultivars.
- Taxon
- winter wheat
Triticum aestivumt0 - Outcome
- grain and biomass yielde0
- Direction
- varied considerablye0
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“effects on grain and biomass yield varied considerably between cultivars”
t0“winter wheat”
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