Abstract
One of the key steps in the oak regeneration process is the successful germination of acorns into new seedlings. Several factors can greatly reduce or entirely destroy a red oak acorn crop between seed fall in the autumn and germination the following spring. In 2001, a bumper acorn crop occurred on Clear Creek State Forest in Jeff erson County, PA. Th is event coincided with the installation of a root development study and the establishment of an oak silviculture short course for SILVAH. Th is fortuitous coincidence allowed for a series of three simple experiments to evaluate factors aff ecting acorn overwintering success and subsequent seedling survival. Th e crop was estimated at approximately 210,000 acorns per acre for all oak species. Insect infestation of the acorns varied among stands, with infestation rates ranging from 5 to 20 percent. Overwintering success of unburied sound acorns was 20 percent, but burial increased survival to more than 80 percent. Unburied acorns succumbed to desiccation (48 percent), insect infestation (15 percent), disease (15 percent), and consumption by wildlife (2 percent). Seedlings that germinated from acorns outside of deer fences and in dense understory shade had the lowest survival rate (2 percent) during the next 8 years. Excluding the deer or reducing the dense understory shade increased survival rates to approximately 26 percent during the same period. Seedlings in fenced, partial-shade stands had an 8-year survival rate of 56 percent. Th ese data indicate that: 1) oak management activities need to be fl exible and opportunistic to take advantage of mast years; 2) there is a several-year window for management following such an event; and 3) dense understory shade and excessive deer browsing are two major obstacles to establishing new oak seedlings, an essential step in the oak regeneration process, at Clear Creek State Forest.
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Claims 11
Claim · model-written summary
The oak acorn crop was estimated at approximately 210,000 per acre.
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“Th e crop was estimated at approximately 210,000 acorns per acre for all oak species.”
Claim · model-written summary
Acorn insect infestation ranged from 5 to 20 percent among stands.
Not reportedInteractorInteractor role
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“Insect infestation of the acorns varied among stands, with infestation rates ranging from 5 to 20 percent.”
e1“for all oak species”
Claim · model-written summary
Unburied sound acorns had 20 percent overwintering success.
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“Overwintering success of unburied sound acorns was 20 percent”
e1“for all oak species”
Claim · model-written summary
Burial increased acorn overwintering survival to more than 80 percent.
- Treatment
- buriale0
- Outcome
- survivale0
- Direction
- increasede0
- Value
- more than 80 percente0
- Comparator
- unburied sound acornse2
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“but burial increased survival to more than 80 percent.”
e1“for all oak species”
e2“Overwintering success of unburied sound acorns was 20 percent”
Claim · model-written summary
Desiccation accounted for 48 percent of unburied acorn losses.
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“Unburied acorns succumbed to desiccation (48 percent)”
e1“for all oak species”
Claim · model-written summary
Insect infestation accounted for 15 percent of unburied acorn losses.
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“insect infestation (15 percent)”
e1“for all oak species”
Claim · model-written summary
Disease accounted for 15 percent of unburied acorn losses.
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“disease (15 percent)”
e1“for all oak species”
Claim · model-written summary
Wildlife consumption accounted for 2 percent of unburied acorn losses.
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“consumption by wildlife (2 percent)”
e1“for all oak species”
Claim · model-written summary
Seedlings outside deer fences in dense shade had 2 percent survival over eight years.
- Outcome
- survival ratee0
- Value
- 2 percente0
- Qualifier
- outside of deer fences and in dense understory shadee0
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“Seedlings that germinated from acorns outside of deer fences and in dense understory shade had the lowest survival rate (2 percent) during the next 8 years.”
e1“for all oak species”
Claim · model-written summary
Deer exclusion or shade reduction increased seedling survival to approximately 26 percent.
- Treatment
- Excluding the deer or reducing the dense understory shadee0
- Outcome
- survival ratese0
- Direction
- increasede0
- Value
- approximately 26 percente0
Not reportedComparator
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“Excluding the deer or reducing the dense understory shade increased survival rates to approximately 26 percent during the same period.”
e1“for all oak species”
Claim · model-written summary
Seedlings in fenced, partial-shade stands had 56 percent survival after eight years.
Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Verbatim
e0“Seedlings in fenced, partial-shade stands had an 8-year survival rate of 56 percent.”
e1“for all oak species”